A secret laboratory near the Arctic Circle is stolen by a mysterious vehicle using a giant magnet. The Anti-Villain League (AVL) tries to recruit former super-villain Gru, now a devoted father to Margo, Edith and Agnes, to find out which evil person stole the lab, which contained a powerful mutagen
known as PX-41 which can make indestructible and extremely aggressive
monsters out of living organisms. They asked Gru because he, being an
ex-villain, knows how villains work.
But Gru refuses to help, saying he
is a legitimate businessman now. Then Gru learns that Dr. Nefario, his
friend and assistant, has decided to leave him for new employment,
because he "missed being evil." Gru reluctantly partners with undercover
AVL agent Lucy Wilde, and together they search The Paradise Shopping
Mall, where they are given a bakery called "Bake My Day" as their
headquarters. Gru suspects Mexican restaurant owner Eduardo Perez of
being a super-villain called "El Macho", a bad guy who supposedly died
after being TNT-laden, skysurfing
a shark into the center of an active volcano. Gru and Lucy break into
Eduardo's restaurant at night, but only find a jar of secret-recipe
salsa and nothing to prove that Eduardo is the culprit. Meanwhile, Agnes
expresses her wish of having a mother someday and believes that Gru
will fall in love with Lucy, despite that Gru tells her that his
relationship with Lucy is strictly professional.
Gru and Lucy investigate wig merchant Floyd Eagle-san, but Gru is
still suspicious of Eduardo. After the investigation, Gru is set up on a
horrible dinner date with a woman named Shannon, who notices Gru's wig
and threatens to humiliate him. Luckily, Lucy bumps into them and
rescues him from the date by shooting Shannon with a mild moose
tranquilizer dart. They take Shannon home, and after they say good
night, Gru realizes that Agnes was right: he has fallen in love with
Lucy.
The next day, the AVL arrests Floyd Eagle-san (who claims that he was
framed) because an almost-empty mutagen jar was found in his shop. The
investigation is closed, and Lucy is reassigned to Australia.
Before leaving, Lucy gives Gru her lipstick taser to remind him of her.
This leaves Gru heartbroken because he loves her, but he cannot find
the courage to ask her out on a date. Instead he brings the girls to
Eduardo's Cinco de Mayo party and finds proof that the Mexican
restaurateur is in fact the super-villain El Macho. Gru discovers that
he (and his helper, who turns out to be Dr. Nefario) have captured and
mutated a large number of Gru's minions using the PX-41, turning them
into insane, savage purple-furred monsters. El Macho plans to send
rockets full of mutated Minions to major cities and achieve world
domination. El Macho emphasizes that he and Gru could team up and they
would become evil again followed by ruling the city, but Gru avoids
answering and leaves with his daughter Margo, who in the meantime has
broken up with Eduardo's two-timing son, Antonio, who had left Margo for
another girl. In his vengeance, Gru uses his freeze-ray to freeze
Antonio in a block of ice.
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