Eep (Emma Stone) is a girl in a family of cavemen living and hunting in pre-historic times. Her family is one of the few to survive, mainly due to the strict rules of her overprotective father, Grug (Nicolas Cage). In their cave home, Grug tells a story to the family, which includes his wife Ugga (Catherine Keener), his daughter Sandy, his son Thunk (Clark Duke), and his mother-in-law Gran (Cloris Leachman). He uses the story of a character who mirrors Eep's curious nature to warn the family that exploration and 'new things' pose a threat to their survival, and says to never not be afraid. This irritates the bored and adventurous Eep, and after the family falls asleep, she leaves the cave, against her father's advice, when she sees a light moving outside.
Seeking the light's source, she meets Guy (Ryan Reynolds), a clever and inventive caveboy. She at first attacks him but then becomes fascinated with the fire he creates and is eager to learn more. He tells her about his theory that the world is reaching its 'end' and asks her to join him. She refuses and Guy leaves, but not before giving her a noise-making shell to call him if she needs help. Eep is then caught by Grug (who had been searching for her), and is later grounded for what she had done. Grug brings Eep home and is joined by the rest of the family. Eep tells them about Guy and shows them the shell given to her, only for them to destroy it in fear of 'new things'. An earthquake then occurs, sending everyone running for the cave, only to be stopped by Grug moments before the cave is destroyed by falling rocks. They climb over the wreckage to discover a land with lush vegetation, much different from their usual surroundings of rocky terrain. Grug takes his family into the forest to find a new cave.
The family is chased by a "Macawnivore" (a large, macaw-colored machairodont later called "Chunky") and attacked by a swarm of "Piranhakeets" (deadly red-feathered, piranha-like birds). In panic, Eep finds and sounds a horn similar to that which Guy gave her. Guy hears this and rushes to her. Thinking quickly, he creates a torch of fire, which scares the birds away. The other Croods are captivated by the fire, having never seen it before. They steal Guy's torch and accidentally set the land around them in flames. Some giant corn is also lit, which rockets up to the sky, prompting a display of "fireworks" as the kernels explode. After feeling impressed by Guy's intelligence and 'ideas', Grug bottles him in a hollow log to carry him in, then suggests that they take solitude in the cave of a nearby mountain mentioned by Guy. Guy is forcibly persuaded to lead the way and learns of the Croods' way of living, which he thinks of as unusual.
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