The spirit of winter Jack Frost is lifted with amnesia from the depths of a frozen pond by the Man in the Moon, only to discover no one can see, hear, or touch him. Three hundred years later, at the North Pole, North, better known as Santa Claus, becomes aware that Pitch Black, the Boogeyman, has returned and is threatening the children of the world. After alerting fellow Guardians the Tooth Fairy (Tooth for short), the Easter Bunny, and the Sandman/Sandy
of the problem, North learns that Jack has been selected by the Man in
the Moon as the newest Guardian. Jack, frustrated and hurt by centuries
of isolation, declines the invitation, but North persuades him to
cooperate by explaining their mission and the looming threat.
Meanwhile, Pitch and his Nightmare minions raid Tooth's palace,
taking all the children's teeth and the memories stored within. He also
captures Tooth's helper fairies except for a single fairy named Baby
Tooth who is rescued by Jack. Resentful that he himself is not believed
in, Pitch plans to destroy children's faith in the Guardians and thereby
weaken them by preventing Tooth from doing her work. To avert this, the
Guardians travel the world collecting teeth, and promise Jack that when
they find Pitch, they will help him regain his own baby teeth and
through them the memories of his past life. On the expedition, a young
boy named Jamie Bennett is delighted to discover the Guardians in his
room. He believes in all of them except for Jack, whom he cannot see.
Pitch learns of the Guardians' resistance and attacks Jack and Sandy.
Despite Jack's efforts to save him, Sandy is overwhelmed and seemingly
killed by Pitch.
The dejected Guardians rally by helping Bunnymund prepare his eggs for Easter.
Although slightly complicated by the intrusion of Jamie's sister
Sophie, the preparation goes smoothly. Jack takes Sophie home, but
afterward a mysterious female voice lures him to Pitch's lair where
Tooth's fairies and the teeth are being held. Pitch keeps him distracted
with the promise of his teeth's memories, while Jack's absence allows
the Nightmares to raid Bunnymund's domain, shattering all the eggs, and
destroying the children's belief in the Easter Bunny.
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