Duke (Channing Tatum) becomes the leader of the G.I. Joe unit, which is framed for stealing nuclear warheads from Pakistan by Zartan (Arnold Vosloo), who is impersonating the President of the United States (Jonathan Pryce).
The majority of the unit is subsequently eliminated in a military air
strike with Duke among the casualties. The only survivors are Roadblock (Dwayne Johnson), Flint (D.J. Cotrona) and Lady Jaye (Adrianne Palicki).
Meanwhile, Storm Shadow (Byung-hun Lee) and ex-Joe Firefly (Ray Stevenson) rescue Cobra Commander (Luke Bracey) from an underground maximum-security prison in Germany, who in turn abandons Destro. Storm Shadow is injured during the escape and retreats to a temple in the Himalayas to recover. Upon learning that he is alive, the Blind Master (RZA), leader of the Arashikage Clan, sends Snake Eyes (Ray Park) and his apprentice Jinx (Elodie Yung), Storm Shadow's cousin, to capture Storm Shadow so he can answer for the murder of his uncle the Hard Master.
Roadblock, Flint, and Lady Jaye return to the United States
where they set up a base of operations in a rundown gym. After Zartan
announces that Cobra will replace the Joes as America's main protective
unit, Lady Jaye deduces that someone is impersonating the President, and
Roadblock leads them to General Joseph Colton (Bruce Willis),
who provides them with weapons, and helps them infiltrate a fundraising
event that the President will be attending. Lady Jaye steals a sample
of the President's DNA and confirms that he is Zartan. They escape after a brief confrontation with Firefly and Zandar (Matt Gerald), the head of the Presidential Detail and a member of Cobra.
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